- SMAD family member 5 (SMAD5)
- SMAD family member 3 (SMAD3)
- Mothers Against Decapentaplegic Homolog 2 (Smad2)
- Cerberus (CER)
- Chordin (CHRD)
- Decorin (DCN)
- Gremlin 1 (GREM1)
- Latent Transforming Growth Factor Beta Binding Protein 1 (LTBP1)
- Noggin (NOG)
- Collagen Type XXVII (COL27)
- Collagen Type XXV (COL25)
- Collagen Type XIX (COL19)
- Collagen Type XVIII (COL18)
- Collagen Type XVII (COL17)
- Collagen Type XVI (COL16)
